Badenian (Middle Miocene) echinoids and starfish from western Ukraine, and their biogeographic and stratigraphic significance

Czasopismo : Acta Geologica Polonica
Tytuł artykułu : Badenian (Middle Miocene) echinoids and starfish from western Ukraine, and their biogeographic and stratigraphic significance

Autorzy :
Thomas, B. A.
Institute of Biological, Environmental and Rural Sciences, Aberystwyth University, Aberystwyth, Ceredigion SY231NP, UK, bat@aber.ac.uk,
Bek, J.
Laboratory of Palaeobiology and Palaeocology, Institute of Geology, Academy of Sciences v.v.i., Rozvojová 269, 165 00 Prague 6, Czech Republic, mrbean@gli.cas.cz,
Hoşgör, I.
Viking International, Şehit Ersan Cad, 24-7, Çankaya-Ankara, Turkey, Izzet.Hosgor@viking-intl.com,
Štamberg, S.
Faculty of Arts, University Hradec Králové, Centre interdisciplinary research, Rokitanského 62, CZ 500 03 Hradec Králové, Czech Republic, s.stamberg@muzeumhk.cz,
Cavin, L.
Department of Geology and Palaeontology, Natural History Museum, CP 6434, 1211 Geneva 6, Switzerland, lionel.cavin@ville-ge.ch,
Grădinaru, E.
Department of Geology, Faculty of Geology and Geophysics, University of Bucharest, Blvd. Bălcescu Nicolae 1, RO-010041 Bucharest, Romania, egradin@geo.edu.ro,
Popa, L.
Department of Geology, Faculty of Geology and Geophysics, University of Bucharest, Bd. Bălcescu Nicolae 1, RO-010041 Bucharest, Romania., livia.dobre@gmail.com,
Panaiotu, C.
Department of Mineralogy, Faculty of Geology and Geophysics, University of Bucharest, Bd. Bălcescu Nicolae 1, RO-010041 Bucharest, Romania, cris.panaiotu@gmail.com,
Grădinaru, E.
Department of Geology, Faculty of Geology and Geophysics, University of Bucharest, Bd. Bălcescu Nicolae 1, RO-010041 Bucharest, Romania, eugen.gradinaru@g.unibuc.ro,
Radwański, A.
Faculty of Geology, University of Warsaw, Al. Żwirki i Wigury 93, PL-02-089 Warszawa, Poland,
Górka, M.
Faculty of Geology, University of Warsaw, Al. Żwirki i Wigury 93, PL-02-089 Warszawa, Poland, magurka@uw.edu.pl,
Wysocka, A.
Faculty of Geology, University of Warsaw, Al. Żwirki i Wigury 93, PL-02-089 Warszawa, Poland,
Abstrakty : Echinoderms from the Badenian (Middle Miocene) of the Fore-Carpathian Basin of western Ukraine are facies restricted. The Mykolaiv Beds, stratigraphically older, yielded the starfish Astropecten forbesi (complete skeletons), two genera of sand dollars (Parascutella, Parmulechinus), and numerous other echinoids of the genera Psammechinus , Echinocyamus, Spatangus, Hemipatagus, Echinocardium, Clypeaster, Echinolampas, and Conolampas. The stratigraphically younger, calcareous Ternopil Beds yielded Eucidaris (complete coronae, isolated spines), Arbacina , Brissus, and Rhabdobrissus. Sixteen species of echinoids are distinguished and/or commented. A new brissid, Rhabdobrissus tarnopolensis sp. nov., is established. A mass occurrence of some species (Psammechinus dubius and Hemipatagus ocellatus) contrasts with that of mass aggregations (sand dollars and Echinocardium leopolitanum) by dynamic events in selected layers of proximal tempestites. Of special note is the occurrence of very small specimens, interpreted as juveniles (‘babies’) having been swept out of their restricted biotopes (‘nurseries’). Some species hitherto regarded as of Early Miocene age, and the problem of their persistence beyond the Fore-Carpathian Basin and/or migration into that basin during the Middle Miocene transgression are discussed.

Słowa kluczowe : asteroidy, jeżowce, taksonomia, nowe gatunki, skamielina, zachowanie, miocen środkowy, baden, Ukraina, asteroids, Echinoids, taxonomy, new species, fossil behaviour, ecotaphonomy, Middle Miocene (Badenian), Ukraine,
Wydawnictwo : Faculty of Geology of the University of Warsaw
Rocznik : 2014
Numer : Vol. 64, no. 2
Strony : 207 – 247
Bibliografia : 1. Agassiz, L. and Desor, E. 1846–1847. Catalogue raisonné des familles, des genres et des espèces de la classe des Échinodermes, p. 32. Paris.
2. Airaghi, C. 1901. Echinidi terziari del Piemonte e della Liguria. Palaeontographia Italica, 7, pp. 149–218.
3. Andreyeva-Grigorovich, A.S., Kulchytsky, Y.O., Gruzman, A.D., Lozynyak, P.Y., Petrashkevich, M.I., Portnyagina, L.O., Ivanina, A.V., Smirnov, S.E., Trofimovich, N.A., Savitskaya, N.A. and Shvareva, N.J. 1997. Regional strati graphic scheme of Neogene formations of the Central Paratethys in the Ukraine. Geologica Carpathica, 48, 123–136.
4. Bałuk, W. and Radwański, A. 1977. Organic communities and facies development of the Korytnica Basin (Middle Miocene; Holy Cross Mountains, Central Poland). Acta Geologica Polonica, 27, 85–123.
5. Blake, D.B. and Zinsmeister, W.J. 1988. Eocene asteroids (Echinodermata) from Seymour Island, Antarctic Peninsula. Memoirs of the Geological Society of America, 169, 489–498.
6. Böhme, M. 2003. The Miocene Climatic Optimum: evidence from ectothermic vertebrates of Central Europe. Palaeogeography Palaeoclimatology Palaeoecology, 195, 389–401.
7. Borghi, E. 1993. Un echinide attuale rinvenuto allo stato fossile: Plagiobrissus (Rhabdobrissus) costai (Gasco, 1876). Bolletino della Societa Reggiana di Scienze Naturali, 13, 1–6.
8. Borghi, E. 1999. Echinodermi fossili dell’Emilia: i Cidaridi del Plio-Pleistocene. Parva Naturalia 1999, pp. 105–126.
9. Boucot, A.J. and Poinar, G.O., Jr. 2010. Fossil Behavior Compendium, pp. 1–391. CRC Press, Taylor & Francis Group; Boca Raton – London – New York.
10. Capeder, G. 1906. Fibularidi del Miocene Medio di S.Gavino a Mare (Portotorres) Sardegna. Bolletino della Societa Geologica Italiana, 25, 495–534.
11. Caso, M.E. 1983. Los equinoideos del Pacifico de México. Parte cuarta: Ordenes Cassiduloida y Spatangoida. Publicaciones especiales, Instituto de Ciencias del Mar y Limnologia, Universidad Nacional Autonoma de México, 6, 1–200.
12. Christensen, A.M. 1970. Feeding biology of the sea-star Astropecten irregularis Pennant. Ophelia, 8, 1–134.
13. Clark, H.L. 1940. Eastern Pacific expeditions of the New York Zoological Society. XXI: Notes on echinoderms from the West Coast of Central America. Zoologica (Scientific Contributions of the New York Zoological Society), 25 (3, Number 22), 331–352.
14. Clark, H.L. 1948. A report on the echini of the warmer Eastern Pacific, based on the collections of the Velero III. Allan Hancock Pacific Expeditions, 8 (5), 225–360. The University of Southern California Press; Los Angeles.
15. Clegg, E.L.G. 1933. Echinoidea from the Persian Gulf. Memoirs of the Geological Survey of India: Palaeontologia Indica, New Series, 22, 1–35.
16. Dembińska-Różkowska, M. 1932. Polnische Miozänkorallen. Annales de la Société Geologique de Pologne, 8, 97–171.
17. Dermitzakis, M.D., Radwańska, U., Radwański, A. and Drina, H. 2009. Lower Pliocene (Zanclean) regressive sequence of Rafina near Pikermi in Attica, Greece: A spectacular locality of mass-aggregated giant balanid cirripedes. Hellenic Journal of Geosciences, 44, 9–19.
18. Desor, E. 1855. Synopsis des Echinides fossiles, p. 38 and pl. 7. Paris – Wiesbade, 1855–1858.
19. Durham, J.W. 1966a. Echinoids: Ecology and Paleontology. In: Moore, R.C. (Ed.), Treatise on Invertebrate Paleontology, Part U (Echinodermata 3, Vol. 1), pp. U211 and U257–U265.
20. Durham, J.W. 1966b. Clypeasteroids. In: Moore, R.C. (Ed.), Treatise on Invertebrate Paleontology, Part U (Echinodermata 3, Vol. 2), pp. U450–U491.
21. Fell, H.B. and Pawson, D.L. 1966. Echinacea. In: Moore, R.C. (Ed.), Treatise on Invertebrate Paleontology, Part U (Echinodermata 3, Vol. 2), pp. U367–U440.
22. Fischer, A.G. 1966. Spatangoids. In: Moore, R.C. (Ed.), Treatise on Invertebrate Paleontology, Part U (Echinodermata 3, Vol. 2), pp. U543–U628.
23. Fourtau, R. 1914. Notes sur les Echinides fossiles de l’Egypte. Bulletin de l’Institut Égyptien. 5 Série, 7 (for 1913), pp. 62–71 and 86–94.
24. Gordon, C.M. and Donovan, S.K. 1992. Disarticulated echinoid ossicles in paleoecology and taphonomy: The last interglacial Falmouth Formation of Jamaica. Palaios, 7, 157–166.
25. Górka, M., Studencka, B., Jasionowski, M., Hara, U., Wysocka, A. and Poberezhskyy, A. 2012. The Medobory Hills (Ukraine): Middle Miocene reef systems in the Paratethys, their biological diversity and lithofacies. Biuletyn Państwowego Instytutu Geologicznego, 449, 147–174.
26. Greenstein, B.J. 1991. An integrated study of echinoid taphonomy: Predictions for the fossil record of four echinoid families. Palaios, 6, 519–540.
27. Hall, K.R.L. and Schaller, G.B. 1964. Tool-using behavior of the California sea otter. Journal of Mammology, 45, 287–298.
28. Harzhauser, M., Mandic, O. and Zuschin, M. 2003. Changes in Paratethyan marine molluscs at the Early/Middle Miocene transition: diversity, palaeogeography and palaeoclimate. Acta Geologica Polonica, 53, 323–339.
29. Heller, C. 1858. Über neue fossile Stelleriden. Sitzungsberichte der Mathematisch-Naturwissenschaftlichen Classe der Kaiserlichen Akademie der Wissenschaften, 28, 155–170.
30. Hendler, G., Miller, J.E., Pawson, D.L. and Kier, P.M. 1995. Sea Stars, Sea Urchins, and Allies: Echinoderms of Florida and the Caribbean. Smithsonian Institution Press, pp. 1–390. Washington – London.
31. Hilber, V. 1882. Geologische Studien in den ostgalizischen Miocän-Gebieten. Jahrbuch der Kaiserlich Königlichen Geologischen Reichsanstalt, 32, 193–330.
32. Jagt, J.W.M. and Codrea, V. 2010. A goniasterid starfish (Echinodermata, Asteroidea) preserved in a mid-Miocene rhyolitic ignimbrite, northwest Romania. Acta Geologica Polonica, 60, 275–282.
33. Jagt, J.W.M. and Ham, R.W.J.M. van der 1994. Early Palaeocene marsupiate regular echinoids from NE Belgium. Proceedings of the Eighth International Echinoderm Conference Dijon/France/6-10 September 1993, pp. 725–729. Balkema; Rotterdam.
34. Janakevich, A.N. 1977. Middle Miocene reefs of Moldavia, pp. 1-116. Stiinca; Kishinev. In Russian.
35. Jankowski, L., Kopciowski, R. and Ryłko, W. 2012. The state of knowledge of geological structures of the Outer Carpathians between Biała and Risca rivers – discussion. Biuletyn Państwowego Instytutu Geologicznego, 449, 203–216. In Polish.
36. Jasionowski, M., Gorka, M., Studencka, B. and Poberezhskyy, A. 2005. Upper Badenian (Middle Miocene) coralline algal reefs of the Medobory Hills (Paratethys, West Ukraine): preliminary results of palaeontological and sedimentological investigations. 12th Congress R.C.M.N.S. 6–11 September 2005, Vienna. Patterns and Processes in the Neogene of the Mediterranean Region. Abstracts, pp. 113–115. Wien.
37. Jasionowski, M., Górka, M., Studencka, B. and Poberezhskyy, A. 2006. Miocene of Medobory Hills Podilya, West Ukraine. In: Wysocka, A. and Jasionowski, M. (Eds), Przebieg i zmienność sedymentacji w basenach przedgórskich. II Polska Konferencja Sedymentologiczna POKOS2, 20-23.06.2006, Zwierzyniec. Przewodnik sesji terenowych, streszczenia referatów i posterów, pp. 53–65. Grodzisk Mazowiecki.
38. Kier, P.M. 1967. Sexual dimorphism in an Eocene echinoid. Journal of Paleontology, 41 (4), 988–993.
39. Kier, P.M. 1969. Sexual dimorphism in fossil echinoids. In: Westermann, G.E.G. (Ed.), Sexual dimorphism in fossil Metazoa and taxonomic implications, pp. 215–222. E. Schweizerbartsche Verlagsbuchhandlung; Stuttgart.
40. Kier, P.M. 1972a. Tertiary and Mesozoic echinoids of Saudi Arabia. Smithsonian Contributions to Paleobiology, 10, 1–242.
41. Kier, P.M. 1972b. Upper Miocene echinoids from the Yorktown Formation of Virginia and their environmental significance. Smithsonian Contributions to Paleobiology, 13, 1–41.
42. Kier, P.M. 1975. The echinoids of Carrie Bow Cay, Belize. Smithsonian Contributions to Zoology, 206, 1–45.
43. Koehler, R. 1915. Echinoderma I: Asteroidea, Ophiuroidea et Echinoidea. In: Michaelsen, W. (Ed.), Beiträge zur Kenntnis der Meeresfauna Westafrikas, Band 1. L. Friederichsen & Co; Hamburg.
44. Kotchetoff, B., Kotchetoff, Y. and Da Veiga Ferreira, O. 1975. Contribution a la connaissance des gisements fossiliferes miocenes au Nord du Cap d’Espichel. Comunicaçoes dos Serviços Geológicos de Portugal, 59, 59–106.
45. Kotlarczyk, J., Jerzmańska, A., Świdnicka, E. and Wiszniowska, T. 2006. A framework of ichthyofaunal ecostratigraphy of the Oligocene – Early Miocene strata of the Polish Outer Carpathian Basin. Annales Societatis Geologorum Poloniae, 76, 1–111.
46. Kroh, A. 2003. The Echinodermata of the Langhian (Lower Badenian) of the Molasse Zone and the northern Vienna Basin (Austria). Annalen des Naturhistorischen Museums in Wien, 104 A, 155–183.
47. Kroh, A. 2005. Echinoidea neogenica. Catalogus Fossilium Austriae, Vol. 2, pp. 1–210. Österreichische Akademie der Wissenschaften; Wien.
48. Kroh, A. 2007a. Hemipatagus, a misinterpreted Loveniid (Echinodermata: Echinoidea). Journal of Systematic Palaeontology, 5, 163–192.
49. Kroh, A. 2007b. Climate changes in the Early to Middle Miocene of the Central Paratethys and the origin of its echinoderm fauna. Palaeogeography, Palaeoclimatology, Palaeoecology, 253, 169–207.
50. Kroh, A. and Harzhauser, M. 1999. An echinoderm fauna from the Lower Miocene of Austria: Paleoecology and implications for Central Paratethys paleobiogeography. Annalen des Naturhistorischen Museums in Wien, 101 A, 145–191.
51. Kroh, A. and Nebelsick, J.H. 2003. Echinoid assemblages as a tool for palaeoenvironmental reconstruction – an example from the Early Miocene of Egypt. Palaeogeography, Palaeoclimatology, Palaeoecology, 201, 157–177.
52. Kroh, A. and Smith, A.B. 2010. The phylogeny and classification of post-Palaeozoic echinoids. Journal of Systematic Palaeontology, 8, 147–212.
53. Kudrin, L.N. 1957. Palaeoecological investigations of the lowermost Tortonian deposits from the south-western part of the Russian Platform. Geologicheskij Sbornik Lvovskovo Gologicheskogo Obshestva, 4, 277–294. In Russian.
54. Lachkhem, H. and Roman, J. 1995. Les Échinoides irréguliers (Néognathostomes et Spatangoides) du Messinien de Melilla (Maroc Septentrional). Annales de Paléontologie (Vert.-Invert.), 81, 247–278.
55. Lambert, J. 1910-1915. Description des Echinides des terrains Néogènes du Bassin du Rhone. Mémoires de la Société Paléontologique Suisse, 37 (1910–1911), 38 (1911–1912), 39 (1913), 41 (1915), pp. 1–240.
56. Lambert, J. 1923. Echinides vivants et fossiles recueillis à Madagascar par M. Raymond Decary. Memoires de la Societe de l'Academie d'Agriculture et des Sciences du Departement de l'Aube, 85, 13–55. Troyes
57. Lambert, J. 1933. Echinides de Madagascar communiques par M.H. Besairie. Annales Géologiques du Service des Mines de Madagascar, 3, 1–49.
58. Laube, G.C. 1871. Die Echinoiden der oesterreichisch-ungarischen oberen Tertiaerablagerungen. Abhandlungen der kaiserlich-königlichen geologischen Reichsanstalt, 5, 55–74.
59. Leeuw, A. de, Bukowski, K., Krijgsman, W. and Kuiper, K.F. 2010. Age of the Badenian salinity crisis; impact of Miocene climate variability on the circum-Mediterranean region. Geology, 38, 715–718.
60. Łomnicki, A.M. 1897. Atlas Geologiczny Galicyi; Tekst do zeszytu dziesiątego, Część 1: Geologia Lwowa i okolicy, pp. 1–208. Komisja Fizyograficzna Akademii Umiejętności; Krakow. In Polish.
61. Łomnicki, A.M. 1898. Atlas Geologiczny Galicyi; Tekst do zeszytu dziesiątego, Część 2, pp. 1–167. Komisya Fizyograficzna Akademii Umiejętności; Krakow. In Polish.
62. Madeira, P., Kroh, A., Cordeiro, R., Meireles, R. and Ávila, S.P. 2011. The fossil echinoids of Santa Maria Island, Azores (Northern Atlantic Ocean). Acta Geologica Polonica, 61, 243–264.
63. McNamara, K.J. 1994. Diversity of Cenozoic marsupiate echinoids as an environmental indicator. Lethaia, 27, 257–268.
64. Mikuž, V. 2009. Scutellids from Miocene beds in Slovenia. Geologija, 52, 205–212.
65. Mortensen, T. 1913. Die Echiniden des Mittelmeeres, eine revidierte Übersicht der im Mittelmeere lebenden Echiniden, mit Bemerkungen über neue oder weniger bekannte Formen. Mitteilungen aus der Zoologischen Station zu Neapel , 21, 1–40.
66. Mortensen, T. 1928. A monograph of the Echinoidea; 1. Cidaroidea, pp. 1–551. C.A. Reitzel, Copenhagen – H. Milford, London.
67. Mortensen, T. 1951. A monograph of the Echinoidea; 5 (2). Spatangoida 2, pp. 1–593. C.A. Reitzel; Copenhagen.
68. Nebelsick, J.H. 1992. The Northern Bay of Safaga (Red Sea, Egypt): an actuopalaeontological approach; III, Distribution of echinoids. Beitrage zur Palaontologie von Osterreich , 17, 5–79.
69. Nebelsick, J.H. 1995. Actuopalaeontological investigations on echinoids: The potential for taphonomic interpretation. In: Emson, R., Smith, A.B. and Campbell, A. (Eds), Echinoderm Research 1995. Proceedings of the fourth European echinoderms colloquium, pp. 209–214. Balkema; Rotterdam/Brookfield.
70. Nebelsick, J.H. and Kroh, A. 2002. The stormy path from life to death assemblages: The formation and preservation of mass accumulations of fossil sand dollars. Palaios, 17, 378–393.
71. Neraudeau, D. 1991. Lateral variations of size-frequency distribution in a fossil echinoid community and their palaeoecological significance. Lethaia, 24, 299–309.
72. Nichols, D. 1959. Changes in the Chalk heart-urchin Micraster interpreted in relation to living forms. Philosophical Transactions of the Royal Society of London, B242 (693), 347–437.
73. Pazdro, Z. 1953. Tectonic units of the Opole Minor, and phases of their development. Bulletin of the Geological Survey of Poland (top secret, for Survey Staff only; unnumbered), pp. 1–114. Wydawnictwa Geologiczne; Warszawa. In Polish and Russian.
74. Peryt, T.M. 2006. The beginning, development and termination of the Middle Miocene Badenian salinity crisis in Central Paratethys. Sedimentary Geology, 188-189, 379–396.
75. Philip, G.M. and Foster, R.J. 1971. Marsupiate Tertiary echinoids from south-eastern Australia and their zoogeographic significance. Palaeontology, 14, 666–695.
76. Philippe, M. 1998. Les échinides miocènes du Bassin du Rhône: révision systématique. Nouvelles Archives du Museum d’Histoire Naturelle de Lyon, 36, 1–441.
77. Pisera, A. 1996. Miocene reefs of the Paratethys: a review. In: Franseen, E.K., Esteban, M., Ward, W.C. and Rouchy, J.-M. (Eds), Models for carbonate stratigraphy from Miocene reef complexes of Mediterranean regions. SEPM Concepts in Sedimentology and Paleontology, 5, 97–104. Tulsa, Oklahoma.
78. Radwański, A. 1970. Dependence of rock-borers and burrowers on the environmental conditions within the Tortonian littoral zone of Southern Poland. In: Crimes, T.P. and Harper, J.C. (Eds), Trace Fossils. Geological Journal Special Issue 3, 371–390. Liverpool.
79. Radwański, A. and Wysocka, A. 2001. Mass aggregation of Middle Miocene spine-coated echinoids Echinocardium and their integrated eco-taphonomy. Acta Geologica Polonica, 51, 295–316.
80. Radwański, A. and Wysocka, A. 2004. A farewell to Świniary sequence of mass-aggregated, spine-coated echinoids Psammechinus and their associates (Middle Miocene; Holy Cross Mountains, Central Poland). Acta Geologica Polonica, 54, 381–399.
81. Radwański, A., Górka, M. and Wysocka, A. 2006. Middle Miocene coralgal facies at Maksymivka near Ternopil (Ukraine): A preliminary account. Acta Geologica Polonica, 56, 89–103.
82. Radwański, A., Wysocka, A. and Górka, M. 2011. ‘Entobia balls’ in the Medobory Biohermal Complex (Middle Miocene, Badenian; western Ukraine). Acta Geologica Polonica, 61, 265–276.
83. Radwański, A., Wysocka, A. and Gorka, M. 2012a. Miocene burrows of the Ghost Crab Ocypode and their environmental significance (Mykolaiv Sands, Fore-Carpathian Basin, Ukraine). Acta Geologica Polonica, 62, 217–229.
84. Radwański, A., Wysocka, A. and Gorka, M. 2012b. Mykolaiv Sands in Opole Minor and beyond: sedimentary features and biotic content of Middle Miocene (Badenian) sand shoals of Western Ukraine. Geological Quarterly, 56, 475–492.
85. Ragaini, L. 1997. Echinoid fauna from the Quaternary fring ing reef of Aqaba (Jordan). Palaeontographia Italica, 84, 1–19.
86. Rakusz, J. 1927. Asteroiden der älteren Mediterran-Stufe aus der Umgebung von Salgotarjan. Földtani Közlöny, 56, 191–195.
87. Roemer, F. 1870. Geologie von Oberschlesien, pp. 1–587. Nischkowsky; Breslau Wrocław.
88. Roman, J. 1970. Description des Échinides. In: C. Montenat and J. Roman, Échinides Néogènes d’Espagne (Provinces d’Alicante et de Murcie). Annales de Paléontologie, Invertebres , 56, 89–138.
89. Roman, J. 1983. Échinides “marsupiaux” (fam. Temnopleuridae) dans le Néogène de l’Ouest européen. Annales de Paleontologie (Vert.-Invert.), 69, 13–42.
90. Sacco, F. 1892. Sopra alcuni Asteroidei fossili. Atti della Reale Accademia delle Scienze di Torino, 25, 739–745.
91. Schneider, S., Berning, B., Bitner, M.A., Carriol, R-P., Jäger, M., Kriwet, J., Kroh, A. and Werner, J. 2009. A parautochthonous shallow marine fauna from the Late Burdigalian (early Ottnangian) of Gurlarn (Lower Bavaria, SE Germany): Macrofaunal inventory and paleoecology. Neues Jahrbuch für Geologie und Palaontologie, Abhandlungen, 254, 63–103.
92. Schultz, H. 2005. Sea urchins; a guide to worldwide shallow water species, pp. 1–484. Partner Scientific Publications; Hemdingen.
93. Seilacher, A. 1979. Constructional morphology of sand dollars. Paleobiology, 5, 191–221.
94. Sismonda, E. 1842. Appendice alla monografia degli Echinidi fossili del Piemonte. Memoria della Reale Accademia della Scienze di Torino, Serie 2, 4, 385–394.
95. Smith, A.B. 1984. Echinoid palaeobiology, pp. 1–190. Allen & Unwin; London.
96. Smith, A.B., Stockley, B. and Godfrey, D. 2010. Spatangoida. In: Smith, A.B. (Ed.), The Echinoid Directory. World Wide Web electronic publication http://www.nhm.ac.uk /palaeontology/echinoids.
97. Szörényi, E. 1953. Miozäne Echinoiden aus den westlichen Teilen der Ukraine. Geologica Hungarica, Series Palaeontologica , 23, 1–122.
98. Śliwiński, M., Bąbel, M., Nejbert, K., Olszewska-Nejbert, D., Gąsiewicz, A., Schreiber, B.C., Benowitz, J.A. and Layer, P. 2012. Badenian–Sarmatian chronostratigraphy in the Polish Carpathian Foredeep. Palaeogeography, Palaeoclimatology, Palaeoecology, 326–328, 12–29.
99. Teisseyre, H. 1938. Quelques observations sur la paléogéographie de Roztocze aux environs de Lwów. Annales de la Société Géologique de Pologne, 14, 113–154.
100. Vadász, M.E. 1915. Die mediterranen Echinodermen Ungarns. Geologica Hungarica, 1 (2), 1–178 (= 79–254).
101. Wiedl, T., Harzhauser, M., Kroh, A., Ćorić, S. and Piller, W.E. 2013. Ecospace variability along a carbonate platform at the northern boundary of the Miocene reef belt (Upper Langhian, Austria). Palaeogeography, Palaeoclimatology, Palaeoecology, http://dx.doi.org/10.1016/j.palaeo.2012.12.015
102. Wright, T. 1855. On fossil echinoderms from the Island of Malta; with notes on the stratigraphical distribution of the fossil organisms in the Maltese beds. Annals and Magazine of Natural History, Series 2, 15 (No. 86), pp. 101–-127, 175–196 and 262–277.
103. Wysocka, A. 1999. Sedymentacja klastycznych utworow badeńskich wschodniej części Roztocza, pomiędzy Zwierzyńcem a Lwowem. Unpublished Ph.D. thesis; Institute of Geology, University of Warsaw. In Polish.
104. Wysocka, A. 2002. Clastic Badenian deposits and sedimentary environments of the Roztocze Hills across the Polish-Ukrainian border. Acta Geologica Polonica, 52, 535–561.
105. Wysocka, A., Radwański, A. and Górka, M. 2012. Mykolaiv Sands in Opole Minor and beyond: Sedimentary features and biotic content of Middle Miocene (Badenian) sand shoals of Western Ukraine. Geological Quarterly, 56, 475–492.
106. Zágoršek, K., Radwańska, U. and Radwański, A. 2012. Bryozoa from the Korytnica Basin (Middle Miocene; Holy Cross Mountains, Central Poland). Bulletin of Geosciences, 87, 201–218.
107. Zapfe, H. 1971. Index Palaeontologicorum Austriae. Catalogus Fossilium Austriae, 15, pp. 1–140. Springer-Verlag; Wien – New York.
DOI :
Cytuj : Thomas, B. A. ,Bek, J. ,Hoşgör, I. ,Štamberg, S. ,Cavin, L. ,Grădinaru, E. ,Popa, L. ,Panaiotu, C. ,Grădinaru, E. ,Radwański, A. ,Górka, M. ,Wysocka, A. , Badenian (Middle Miocene) echinoids and starfish from western Ukraine, and their biogeographic and stratigraphic significance. Acta Geologica Polonica Vol. 64, no. 2/2014
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